• B.Sc. (Hons.), Botany, University of Karachi, 1973 
• M.Sc., Plant Physiology, University of Karachi, 1974 
• Ph.D., Physiological Ecology, Ohio University, 1985 
• D.Sc., Botany, University of Karachi, 2010.

Salt tolerant plants (halophytes), physiology, biochemistry and ecophysiology; Cash-crop halophytes

Development of varieties of halophytes for special uses
Salt tolerance mechanisms
Seed biology of halophytes 

Physiological Ecology of Plants
Plant Ecology
Plant Population Biology
Seed Biology
Evolutionary Ecology

• Koyro, H-W, M. A. Khan and H. Lieth. 2011. Halophytic crops: A resource for the future to reduce the water crisis? Emir. J. Food Agric. 2011. 23: 001-016
• Abideen, Z., R. Ansari and M. A. Khan. 2011. Halophytes: Potential source of ligno-cellulosic biomass for ethanol production. Biomass & Bioenergy. 35: 1818-1822.
• Qadir, M., A. Tubeileh, J. Akhtar, A. Larbi, P.S. Minhas, and M.A. Khan. 2008. Productivity enhancement of salt-prone land and water resources through crop diversification. Land Degradation and Development. 19: 429-453.
• Siddiqui, Z.S., M.A. Khan, B-G. Kim, J-S. Huang, T-R. Kwon. 2008. Physiological response of Brassica napus genotypes to combined drought and salt stress. Plant Stress 2: 78 - 83.
• Weber, D. J., R. Ansari, B. Gul, and M. A. Khan. 2006. Potential of halophytes as source of edible oil. Journal of Arid Environment 68: 315-321.
• Khan, M.A., R. Ansari, H. Ali, B. Gul and B. L. Nielsen. 2009. Panicum turgidum: a sustainable feed alternative for cattle in saline areas. Agriculture, Ecosystems & Environment 129: 542-546.
• Khan, M.A., R. Ansari, B. Gul and M. Qadir. 2006. Crop diversification options for salt-prone land resources. In Perspectives in Agriculture, Veterinary Science, Nutrition and Natural Resources. CABI, International, Vol 1 number 48.
• Gulzar, S., M. A. Khan and X. Liu. 2007. Seed germination strategies of Desmostachya bipinnata: a fodder crop for saline soils. Rangeland Ecology and Management 60: 401-407.
• Gulzar, S. and M.A. Khan. 2001. Seed germination of a halophytic grass Aeluropus lagopoides. Annals of Botany 87: 319-324.
• Khan, M.A. and S. Gulzar. 2003. Light, salinity and temperature effects on the seed germination of perennial grasses. American Journal of Botany 90: 131-134.
• Khan, M.A., I.A. Ungar and A.M. Showalter. 2000. Salt tolerance in the subtropical perennial halophyte Atriplex griffithii Moq. var. stocksii Boiss. Annals of Botany 85: 225-232.
• Khan, M.A. and B. Gul. 1998. High salt tolerance in the germinating dimorphic seeds of Arthrocnemum indicum. International Journal of Plant Sciences 159: 826-832.
• Khan, M.A., I.A. Ungar, A.M. Showalter and H. Dewald. 1998. NaCl - induced accumulation of glycinebetaine in four subtropical halophytes from Pakistan. Physiologia Plantarum 102: 487-492.
• Khan, M.A. and I.A. Ungar. 1997. Effect of thermoperiod on recovery of seed germination of halophytes from saline conditions. American Journal of Botany 84: 279-283.
• Khan, M.A. and I.A. Ungar. 1996. Influence of salinity and temperature on the germination of Haloxylon recurvum. Annals of Botany 78: 547-551.
• Khan, M.A. and D.J. Weber. 1986. Factors influencing seed germination in Salicornia pacifica var. utahensis. American Journal of Botany 73: 1163-1167.
• Chrominski, A., M.A. Khan, D.J. Weber and B.N. Smith. 1986. Ethylene and ethane production in response to salinity stress. Plant, Cell & Environment 9: 687-691.
• Khan, M.A. and I.A. Ungar. 1985. The role of hormone in regulating germination of polymorphic seeds and early seedling growth of Atriplex triangularis under saline conditions. Physiologia Plantarum 63: 109-113.
• Khan, M.A. and I.A. Ungar. 1984. Seed polymorphism and germination responses to salinity stress in Atriplex triangularis Willd. Botanical Gazzette 145: 487-494.
• Khan, M.A. and I.A. Ungar. 1984. Effects of salinity and temperature on the germination and growth of Atriplex triangularis Willd. American Journal of Botany. 71: 481-489.